A Machine Gun Company was attached to each Infantry Brigade and their subsequent Division, an additional Company was attached as a divisional reserve.
| Brigade attached to: | Divisional Reserve |
| Division attached to: | 16th (Irish) Division |
The 269th M.G. Coy. disembarked at le Havre on 15 January 1918 and joined the Division on 18 January 1918.
By 09 March 1918, it was amalgamated with the 47th, 48th and 49th Machine Gun Companies to form No. 16 Battalion, Machine Gun Corps.